下列那一種酵素參與克式循環(Krebs cycle,亦稱citric acid cycle或TCA cycle)的反應,是位於粒線體內膜上且直接參與電子傳遞鏈?
succinate dehydrogenase
isocitrate dehydrogenase
alpha-ketoglutarate dehydrogenase complex
malate dehydrogenase