細胞進行糖質新生（gluconeogenesis）時，需要下列那兩種酵素共同作用將pyruvate轉換為phosphoenolpyruvate？
pyruvate kinase and pyruvate dehydrogenase
pyruvate carboxylase and pyruvate kinase
pyruvate carboxylase and phosphoenolpyruvate carboxykinase
pyruvate dehydrogenease and phosphoenolpyruvate carboxykinase
